Education

MD/PhD · 2026 (expected)
University of California, San Francisco

PhD in the UCSF Neuroscience Graduate Division · Weill Fellow

BS · 2015
Johns Hopkins University

Dual major in Biomedical Engineering and Mathematics · General Honors · Departmental Honors (BME, Math) · Richard J. Johns Award (BME outstanding scholarship) · Michael Bloomberg Scholar · Elected to Tau Beta Pi

Research Experience

Poskanzer Lab · 2020–present
University of California, San Francisco

Projects: Astrocyte closed-loop regulation of neuronal dynamics · Astrocytic calcium response to neurotransmitters · Astrocytic GPCR signaling regulating sleep

Crone Lab · 2012–2018
Johns Hopkins University

Projects: Dynamics of information flow during cognition · Networks for language and movement · Tools to connect brain activity to the web · Language brain-machine interfaces

Hong Lab · 2013
Tsinghua University

Projects: Networks for language

Teaching

Presenter · 2021
Neuroscience Orientation · UCSF

Professor: Vijay Namboodiri. Led a two-hour tutorial on probability, statistics, and experimental design.

TA · 2014
Calculus III · Johns Hopkins University

Professor: Richard Brown. Led a section of ~30 students once per week. Content covered multivariate calculus.

Publications

Cortical astrocytes independently regulate sleep depth and duration via separate GPCR pathways
Vaidyanathan, T. V., Collard, M., Yokoyama, S., Reitman, M. E., & Poskanzer, K. E. (2021). Elife, 10, e63329.

Changes in human brain dynamics during behavioral priming and repetition suppression
Korzeniewska, A., Wang, Y., Benz, H. L., Fifer, M. S., Collard, M., Milsap, G., ... & Crone, N. E. (2020). Progress in neurobiology, 189, 101788.

Keyword Spotting Using Human Electrocorticographic Recordings
Milsap, G., Collard, M., Coogan, C., Rabbani, Q., Wang, Y., & Crone, N. E. (2019). Frontiers in neuroscience, 13, 60.

BCI2000Web and WebFM: browser-based tools for brain computer interfaces and functional brain mapping
Milsap, G., Collard, M., Coogan, C., & Crone, N. E. (2019). Frontiers in neuroscience, 12, 1030.

Cortical responses to input from distant areas are modulated by local spontaneous alpha/beta oscillations
Usami, K., Milsap, G. W., Korzeniewska, A., Collard, M. J., Wang, Y., Lesser, R. P., ... & Crone, N. E. (2019). Cerebral cortex, 29(2), 777-787.

Cortical subnetwork dynamics during human language tasks
Collard, M. J., Fifer, M. S., Benz, H. L., McMullen, D. P., Wang, Y., Milsap, G. W., ... & Crone, N. E. (2016). Neuroimage, 135, 261-272.